수목의 (樹木) 수분특성에 관한 생리 · 생태학적 해석 - 물참나무와 상수리나무 엽의 수분특성의 계절변화 -

한상섭,최흥선 ( Sang Sup Han,Heung Seon Choi ) 한국산림과학회 1986 한국산림과학회지 Vol.72 No.1

Seasonal changes of water relations parameters were obtained from p-v curves in leaves of Quercus grosseserrata and Quercus acutissima. The osmotic pressure at full hydration, no, and osmotic pressure at incipient plasmolysis, trp, were high in newly emerged leaves but decreased with leaf development in each of the species. Water deficit at turgor loss was 10 to 20% in each of the species during the growing season. Maximum bulk elastic modulus in cell walls at full turgor, Emax, rises rapidly with leaf development before senescence in each of the species. Seasonal change of number of osmoles solute in symplasm per dry weight, Ns/DW, was higher in Quercus grosseserrata leaves than Quercus acutissima leaves, while relative water content (Vp/Vo, RWC^*, Vo/Vt) was relatively constant in each of the species.

수목(樹木)의 수분특성(水分特性)에 관(關)한 생리(生理)·생태학적(生態學的) 해석(解析)(VIII) - 신갈나무의 수분통도성(水分通導性) 구조(構造) -

한상섭,김선희,Han, Sang Sup,Kim, Sun Hee 한국산림과학회 1996 한국산림과학회지 Vol.85 No.1

본 연구는 우리 나라 대표적인 향토수종의 하나인 신갈나무의 수분 생리적 특성을 밝히고자 하는 일련의 연구로서, 줄기, 가지, 분지부분, 그리고 정단부 1년생 소지에 있어서 상대수분통도성, Leaf specific conductivity(LSC), Huber value, Specific conductivity, 부위별 평균도관직경, 단위면적당 도관수 등을 측정하여 수체내 수분통도성의 공간적 배치구조(Hydraulic architecture)를 비교 고찰하였다. 1. 신갈나무의 상대수분통도성은 줄기에서 $2.5526{\times}10^{-12}{\sim}1.2260{\times}10^{-10}m^2$, 가지에서 $1.6279{\times}10^{-11}{\sim}6.8378{\times}10^{-11}m^2$의 범위로 나타냈다. 상대수분통도성은 줄기와 가지의 직경이 작아질수록 큰 값을 나타냈다. 정단부 1년생 소지에 있어서는 주지가 측지보다 평균 2배 이상 큰 값을 나타냈다. 2. LSC는 줄기 하부에서 작은 값을 나타냈고, 상부로 갈수록 큰 값을 나타냈다. 그리고 줄기가 가지보다 월등히 큰 값을 보였으며, 분지부분이 가장 작은 값을 나타냈다. 3. 분지부분의 수분통도성값은 분지형태(ㅏ형과 Y형)에 따라 차이를 보였다. ㅏ형의 분지부분에서는 상대수분통도성, LSC, Specific conductivity, 평균도관직경 등이 줄기가 가지 보다 2 배 이상 큰 값을 나타냈고, Y형에서는 1~1.5배 더 컸다. 4. 줄기의 LSC, Specific conductivity, Huber value는 직경이 작아질수록 증가하였다. 5. 정단부 1년생 소지에 있어서 LSC, Huber value, 평균도관직경은 주지가 측지보다 큰 값을 나타냈다. 6. 줄기의 부위별 평균도관직경은 줄기의 직경이 작아질수록 감소한 반면, 단위면적당 도관수($mm^{-2}$)는 많아졌다. 줄기에 있어서 도관직경은 직경이 작아질수록 조재부에서 급격하게, 만재부에서는 완만하게 감소하였다. This study was carried out to investigate the hydraulic architecture such as relative hydraulic conductivity, Leaf specific conductivity(LSC), Huber value, Specific conductivity of the stem, branch and Junctions of stem-to-branch in Quercus mongolica trees. The hydraulic architecture of various hydraulic conductivities of stem and branch was described. The results obtained were summarized as follows : 1. The range of relative hydraulic conductivity was $2.5526{\times}10^{-12}$ to $1.2260{\times}10^{-10}m^2$ in stems, $1.6279{\times}10^{-11}$ to $6.8378{\times}10^{-11}m^2$ in branches. The relative hydraulic conductivities increased with decreasing diameter of stem and branch. The relative hydraulic conductivity of one-year-old terminal shoots were two times greater than that of the lateral shoots. 2. LSC value was larger at the top than at the base in stem. LSC is much smaller in branches than in stem ; especially smallest at branching part. 3. Hydraulic conductivities of the branching part appeared the different values with the 4 type and 4 type. Relative hydraulic conductivity, LSC, Specific conductivity and mean vessel diameter in type branching part were larger in stem than in branch part, but not found in the branching part of Y type. 4. LSC and Specific conductivity of stem increased with decreasing diameter, but Huber value slowly increased with decreasing diameter ; especially highest at less than 1cm diameter. 5. LSC, Huber value, and mean diameter of vessels were larger at 1-year-old leader shoots than at lateral shoots. 6. The mean vessel diameter in various parts of a tree decreased with decreasing diameter of stem, but the number of vessels per unit area($mm^{-2}$) increased reversely. Mean vessel diameter in stem decreased sharply at earlywood and slowly at latewood with decreasing diameter of stem.

Studies on the Productivity of Korean White Pine Forest (I) Effects of Temperature, Light and Water Stress on Photosynthesis and Dark Respiration Rates of Leaves

한상섭,Han, Sang Sup Korean Society of Forest Science 1982 한국산림과학회지 Vol.55 No.1

본 연구(硏究)는 잣나무림(林)의 일차(一次) 생산력(生産力)을 알기위한 것으로, 엽(葉)의 온도(溫度) 광도(光度) 수분결핍(水分缺乏) 등이 광합성속도(光合成速度)및 호흡속도(呼吸速度)에 미치는 영향(影響)을 측정(測定) 보고(報告)한다. 1) 광포화(光飽和)는 40Klwx 전후에서 일어나며 광보상점(光補償点)은 l.0~1.3Klwx 임을 알았다. 2) 당년엽(當年葉)의 광합성(光合成) 속도(速度)가 가장 높고 엽령(葉令)이 증가함에 따라 감소(減少)했다. 3) 온도반응(溫度反應)에 따른 호흡속도(呼吸速度)는 2월엽(月葉)이 8월엽(月葉)보다 2배(倍)정도 높았고, 양자(兩者) 모두 $17^{\circ}C$ 이하에서 심한 감소(減少)를 보였다. 4) 광합성(光合成) 속도(速度)는 약 -10bar에서 감소(減少)가 시작되며 약 -24bar에서 0으로 되었다. 호흡속도(呼吸速度)는 약 -5 bar에서 감소(減少)가 시작 -24bar에서 43% 정도 감소했다. 5) 광합성(光合成)에 관한 최적온도(最適溫度)는 8월 엽(葉)에서 $23{\sim}26^{\circ}C$, 2월엽(月葉)에서 $15{\sim}18^{\circ}C$를 나타냈다. This study is to investigate the effects of temperature, light and water deficit on apparent phytosynthesis rate (Pn) and dark respiration rate(Rd) of leaves in the series of studies dealing with primary productivity of korean white pine forest. The results obtained are as follows: 1. The light saturation for Pn occured at about 40 Klux, and light compensation at 1.0 to 1.3 Klux. 2. The Pn of current leaves was highest, and Pn was decreased with increasing leaf age. 3. The Rd on the response of temperature in February was about two times value in all of the temperature ranges as compared with the ones in August. 4. The incipient water stress, above which Pn and Rd declined from 100%, was different for Pn(-10bar). The high water stress required to reduce Pn to nearly 0%, at -24 bar, but Rd was only 43% at -24 bar. 5. The optimum temperature range for Pn showed about 15 to $18^{\circ}C$ in February and 23 to $26^{\circ}C$ in August.

수목(樹木)의 수분특성(水分特性)에 관(關)한 생리(生理)·생태학적(生態學的) 해석(解析)(XII) - 낙엽(落葉) 활엽수(闊葉樹) 21종(種)의 수체(樹體) 부위별(部位別) Leaf Specific Conductivity의 특성(特性) -

한상섭,김선희,Han, Sang Sup,Kim, Sun Hee 한국산림과학회 1999 한국산림과학회지 Vol.88 No.1

이 연구는 우리 나라 주요(主要) 낙엽(落葉) 활엽수(闊葉樹) 21종(種)에 대하여 줄기, 가지, 분지부(分技部)와 정단부(頂端部) 1년생 주간(主幹)과 측지(側枝) 등의 leaf specific conductivity(LSC)를 측정하여 수분통도성(水分通導性)의 배치구조(hydraulic architecture)를 수종별로 비교하였다. 고로쇠나무, 복자기, 박달나무, 물푸레나무와 느티나무를 제외한 활엽수종의 LSC 값은 줄기가 가지보다 컸으며, 분지부(分技部)는 가장 작은 값을 나타냈다. 특히 분지부(分技部)에 있어서 LSC값의 급격한 감소는 모든 수종에서 나타난 공통적인 특징이다. 산공재 수종에서는 자작나무, 피나무, 층층나무, 박달나무가 다른 수종에 비하여 높은 LSC를 나타냈으며, 복자기는 가장 낮은 LSC를 나타내는 수종이었다. 환공재 수종에서는 다릅나무, 참오동나무, 갈참나무, 졸참나무의 LSC가 높았으며, 느릅나무가 가장 낮은 LSC를 나타냈다. 복자기, 층층나무, 들메나무, 신갈나무와 느티나무에 있어서 정단부(頂端部) 1년생 주간(主幹) 및 측지(側枝)의 LSC값은 주간이 최소 $22{\mu}{\ell}/g$, 최대 $139{\mu}{\ell}/g$ 이고, 측지가 최소 $11{\mu}{\ell}/g$, 최대 $73{\mu}{\ell}/g$의 범위를 보였으며, 정단주간(頂端主幹)(leaders)이 정단측지(頂端側技)(adjacent laterals)보다 평균 2배 이상, 최대 5배 정도로 크게 나타나, 정단주간은 정단측지보다 수력적(水力的) 우세(優勢)(hydraulic dominance)를 보였다. This study was carried out to compare the characteristics of hydraulic architecture by measurement of leaf specific conductivity(LSC) in the twenty-one deciduous broad-leaved species. The LSC's of stems were higher than in branches except Acer mono, A. triflorum, Betula schmidtii, Fraxinus rhynchophylla and Zelkova serrata, and lowest in junction parts as compared with stems and branches. In diffuse-porous species, the LSC was much higher in B. plalyphylla var. iaponica, B. schmidtii, Cornus controversa, Tilia amurensis than those of other species and especially lowest in A. triflorum. In ring-porous species, the LSC was much higher in Maackia amurensis, Paulownia tomentasa, Quercus aliens, Q. serrata than those of other species and lowest in Ulmus davidiana var. japonica. The LSC's of 1-year-old terminal shoots of stem(the leader and adjacent laterals) in A. triflorum, C. controversa. F. mandshurica, Q. mongolica and Z. serrata ranged from 22 to $139{\mu}{\ell}/g$ in the leaders, and 11 to $73{\mu}{\ell}/g$ in the adjacent laterals. The LSC's of leaders were usually greater than the adjacent laterals because of the hydraulic dominance of the leader shoot.

수목의 수분특성에 관한 생리 · 생태학적 해석 (Ⅲ) - 몇 종의 침엽수에 있어서 Shoot Water Potential 의 일변화 및 Xylem Conductivity 의 특성 -

한상섭,전두식 ( Sang Sup Han,Doo Sik Jeon ) 한국산림과학회 1984 한국산림과학회지 Vol.63 No.1

This study was to investigate the diurnal changes of shoot water potentials and the characteristics of xylem conductivity of branch in several conifers. The results obtained are as follows: 1 ) The diurnal shoot `water potentials fluctuated with the sunlight intensities, and increase in shoot water potential lagged behind two hours as compared with the time of sunlight decrease in tree crown. 2) The shoot water potential reached the daily maximum ai twelve to fourteen o`clock in the afternoon, and the maximum shoot water potentials were -22 bar in Larix leptolepis, -18 bar in Pinus koraiensis, -15 bar in Pinus densiflora, -14 bar in Abies holophylla, and -10 in Pinus rigida. 3)The average gradient of shoot water potential per one meter height (φ_L/m) in tree crown was -1.7 bar/m in Pinus koraiensis while that of Larix leptolepis was -2.1 bar/m. 4) The average of relative xylem conductivities (K, ㎠/hr.atm) in branches was 2878 in Larix leptolepis, 2763 in Pinus rigida,

수목의 (樹木) 수분특성에 관한 생리 · 생태학적 해석 (Ⅰ) - Pressure Chamber Technique 에 의한 내건성 (耐乾性) 수종의 (樹種) 진단 -

한상섭,김광륜 ( Sang Sup Han,Kwang Ryun Kim ) 한국산림과학회 1980 한국산림과학회지 Vol.50 No.1

The purpose of the this study was to compare the torlent tree to drought among the Pinus koraiensis, Abies holophylla, and Ginkgo biloba. The water relations parameters of leafy shoots were measured by the Pressure chamber technique from August 10 to September 4 in 1980. On the water relations parameters such as original osmotic pressure(π_0), osmotic pressure(π_p) and relative water content (RWC) at incipient plasmolysis, and Vp/Vo ratio of the volume of osmotic water (Vp) at incipient plasmolysis to total symplasmic water (Vo) basis, the Abies holophylla shoots showed ontogenetically a superior osmoregulation which are closely associated with drought resistance compared with Pinus koraiensis, and the Ginkgo biloba shoots showed the heighest among these three species.

잣나무 구과의 (毬果) 광합성과 호흡의 특성에 관한 연구

한상섭,김영모 ( Sang Sup Han,Young Mo Kim ) 한국산림과학회 1988 한국산림과학회지 Vol.77 No.1

The dark respiration, photosynthesis(CO₂ refixation), CO₂ balance and chlorophyll content of lst-year conelets and 2nd-year cones of Korean pine(Pinus koraiensis S.et.Z.) were investigated after pollination up to the end of maturation. The results obtained are as follows : 1. The growth of 1st-year conelet was 3.6㎝ in length. 2.4㎝ in diameter and 3.058 in dry weighs during the first year. The growth of 2nd-year mature cone was 13.5cm in length, 9.3㎝ in diameter and 141.0g in dry weight in the late of 2nd-year. 2. The refixation of carbon dioxide released from a cone by the dark respiration was less than 50 percent at light saturation through the growing period. The refixation of carbon dioxide released by dark respiration for one year was 7.3 percent in 1st-year conelets and 8.7 percent in 2nd-year cones. 3. The dark respiration rate of cones by increasing temperature was rapidly increased with increasing temperature up to 25℃. The dark respiration rate of cones was much higher in non-growing season than that in growing season at the same temperature. 4. The rates of dark respiration and CO₂ refixation, based on the dry weight, were much higher in 1st-year conelet than that in 2nd-year cone. 5. The CO₂ balance for a cone was negative from pollination to the end of maturation. The net dark respiration loss for a cone was 7.23g CO₂/year in 1st-year conelet and 164.8g CO₂/year in 2nd-year cone. 6. The respiratory loss efficiency for a cone(=CH₂O weight calculated by net dark respirationjdry weight of cone) for one year was 1.61 in 1st-year conelet and 0.81 in 2nd-year cone for one year. 7. The total chlorophyll content of surface scale of the cone was lower than 2㎎/g dw through the growing period, and chl. a/b ratio was 2 to 3.

수목(樹木)의 수분특성(水分特性)에 관한 생리(生理)·생태학적(生態學的) 해석(解析(IV) - 몇 종(種)의 침엽수(針葉樹)에 있어서 Leaf Conductance와 Water Potential, 상대함수율(相對含水率), 팽압(膨壓)과의 관계(關係) -

한상섭,전두식,Han, Sang Sup,Jeon, Doo Sik 한국산림과학회 1984 한국산림과학회지 Vol.63 No.1

본연구(本硏究)는 몇 종(種)의 침엽수(針葉樹)의 지엽(枝葉)에 있어서 P-V 곡선법(曲線法)에 의해 얻은 수분특성인자(水分特性因子)와 기공폐쇄작용(氣孔閉?作用)과의 관계(關係)를 측정고찰(測定考察)한 것으로 그 결과(結果)는 다음과 같다. 1. P-V 곡선법(曲線法)에 의해 구(求)한 초기원형질리점(初期原形質離點)의 water potential 값과 완전기공폐쇄점(完全氣孔閉?點)의 water potential 값은 거의 일치(一致)하였다. 2) 완전기공폐쇄점(完全氣孔閉?)는 잣나무 -21 bar(음엽(陰葉) -17 bar), 리기다 -20 bar, 소나무 -22 bar, 일본잎갈나무 -24 bar 전후(前後)에서 일어났다. 상대함수율(相對含水率)로 나타내면 잣나무 85%(음엽(陰葉) 82%), 리기다 77%, 소나무 85%, 일본잎갈나무 70% 전후(前後)에서 완전기공폐쇄(完全氣孔閉?)가 일어났다. 3. 초기기공폐쇄(初期氣孔閉?)는 잣나무 -14 bar(음엽(陰葉) -12 bar), 리기다 -10 bar, 소나무 -15 bar, 일본잎갈나무 -6 bar 전후(前後)에서 일어났다. 상대함수율(相對含水率)로 나타내면 잣나무 90%, 리기다 93%, 소나무 90% 일본잎갈나무 93% 전후(前後)에서 초기기공폐쇄(初期氣孔閉?)가 일어났다. 4. leaf conductance가 증가(增加)함에 따라 팽압(膨壓)은 증가(增加)하며, 상대함수율(相對含水率)이 증가(增加)함에 따라 팽압(膨壓)은 직선적(直線的)으로 증가(增加)하였다. This study was to elucidate the relation between the water relations parameters obtained from P-V curves and stomatal closure. The results obtained are as follows: 1) The water potential at incipient plasmolysis obtained from P-V curves was similar to the water potential at critical stomatal closure. 2) The critical stomatal closure of sun leaves appear at -21 bar (-17 bar, shade leaves) in Pinus koraiensis, -20 bar in Pinus rigida, -22 bar in Pinus densiflora, and -24 bar in Larix leptolepis. On a relative water content basis, the critical stomatal closures of sun leaves appear at 85% (82%, shade leaves) in Pinus kordiensis, 77% in Pinus ragida, 85% in Pinus densiflora, and 70% in Larix leptolepis. 3) The incipient stomatal closures of sun leaves appear at -14 bar (-12 bar, shade leaves) in Pinus koraiensis, -10 bar in Pinus rigida, -15 bar in Pinus densiflora, and -6 bar in Larix leptolepis. On a relative water content basis, the incipient stomatal closures of sun leaves appear at 90% in Pinus koraiensis, 93% in Pinus rigida, 90% in Pinus densiflora, and 93% in Larix leptolepis. 4) The leaf conductance was increased by increase in volume-averaged turgor pressure was linearly increased by increase in relative water content.

수목(樹木)의 수분특성(水分特性)에 관한 생리(生理)·생태학적(生態學的) 해석(解析(III) - 몇 종(種)의 침엽수(針葉樹)에 있어서 Shoot Water Potential의 일변화(日變化) 및 Xylem Conductivity의 특성(特性) -

한상섭,전두식,Han, Sang Sup,Jeon, Doo Sik 한국산림과학회 1984 한국산림과학회지 Vol.63 No.1

본(本) 연구(硏究)는 몇 종(種)의 침엽수(針葉樹)에 대하여 Shoot water potential의 일변화(日變化) 및 지(枝)의 Xylem conductivity의 특성(特性)을 측정고찰(測定考察)한 것으로 다음과 같은 결과를 얻었다. 1) Shoot water potential은 광도(光度)의 크기에 따라 변화(變化)하며, 광도(光度)의 감소(減少)가 시작된지 2시간(時間) 늦게 Shoot water potential의 증가(增加)가 시작되었다. 2) 지엽(枝葉)의 일중(日中) 최대수분(最大水分) 결차(缺差)는 12시(時)부터 14시(時) 사이에 일어나며, 그 값의 크기는 일본잎갈나무 -22 bar, 잣나무 -18 bar, 소나무 -15 bar, 젓나무 -14 bar, 리기다소나무 -10 bar 정도였다. 3) 수고(樹高) 1 m 당(當) 지엽(枝葉)의 수분결차(水分缺差)의 크기를 경사도(傾斜度)(${\varphi}_L/m$)로 나타내면 잣나무 -1.7 bar/m, 일본잎갈나무 -2.1 bar/m였다. 4) 지(枝)의 Relative Xylem conductivity (K, $cm^2/hr{\cdot}atm$)의 평균치(平均値)는 일본잎갈나무 2878, 리기다소나무 2763, 소나무 2652, 잣나무 2113 이었다. This study was to investigate the diurnal changes of shoot water potentials and the characteristics of xylem conductivity of branch in several conifers. The results obtained are as follows: 1) The diurnal shoot water potentials fluctuated with the sunlight intensities, and increase in shoot water potential lagged behind two hours as compared with the time of sunlight decrease in tree crown. 2) The shoot water potential reached the daily maximum ai twelve to fourteen o'clock in the afternoon, and the maximum shoot water potentials were -22 bar in Larix leptolepis, -18 bar in Pinus koraiensis, -15 bar in Pinus densiflora, -14 bar in Abies holophylla, and -10 in Pinus rigida. 3) The average gradient of shoot water potential per one meter height (${\varphi}_L/m$) in tree crown was -1.7 bar/m in Pinus koraiensis while that of Larix leptolepis was -2.1 bar/m. 4) The average of relative xylem conductivities (K, $cm^2/hr{\cdot}atm$) in branches was 2878 in Larix leptolepis, 2763 in Pinus rigida, 2652 in Pinus densiflora, and 2113 in Pinus koraiensis.

수목(樹木)의 수분특성(水分特性)에 관(關)한 생리(生理)·생태학적(生態學的) 해석(解析)(VII) - Heat pulse법(法)에 의한 낙엽송임분(林分)의 수액류속(樹液流速) 계측(計測) -

한상섭,김선희,Han, Sang Sup,Kim, Sun Hee 한국산림과학회 1993 한국산림과학회지 Vol.82 No.2

본(本) 연구(硏究)는 heat pulse법(法)을 이용하여 낙엽송 임분(林分)의 증산량(蒸散量)을 알기 위한 기초 연구로써, 일사량(日射量), 온도(溫度), 습도(濕度) 등의 변화에 따른 heat pulse 속도(速度)의 일변화와 계절변화, 방위별 heat pulse 속도(速度)의 차이, 변재부(邊材部)에 있어서 깊이별 heat pulse 속도(速度)의 차이, 우세목(優勢木), 준우세목(準優勢木), 열세목(劣勢木)의 heat pulse 속도(速度)의 차이, 엽(葉)의 수분(水分)포텐셜과 heat pulse 속도(速度)와의 관계, 줄기에 있어 수분상승의 방향, heat pulse법(法)으로 추산한 임분(林分)의 증산량(蒸散量)등에 대하여 측정 고찰하였다. 얻어진 결과를 요약하연 다음과 같다. 1 낙엽송의 heat pulse 속도(速度)(V)와 수액유속(樹液流速)(SFR)과의 관계는 SFR=1.37V($r=0.96^{**}$)의 식으로 나타냈다. 2. 우세목(優勢木), 준우세목(準優勢木), 열세목(劣勢木)의 heat pulse 속도(速度)를 비교하면, 수액유속(樹液流速)은 우세목(優勢木)이 가장 높고, 준우세목(準優勢木), 열세목(劣勢木) 순위였다. Heat pulse 속도(速度)는 일사량(日射量), 온도(溫度), 대기포차(大氣飽差) 등의 크기에 따라 심한 변화(變化)를 나타냈다. 3. 입목(立木) 개체간(個體間)의 heat pulse 속도(速度)의 차이는 이른 아침과 밤에는 거의 없으나, 일사량(日射量)이 높은 12시부터 16시 사이에는 약간의 차이를 나타냈다. 4. Heat pulse 속도(速度)와 수분(水分)포텐셜은 거의 비슷한 일변화 경향을 나타냈다. 5. Heat pulse 속도(速度)의 계절변화는 8월에 가장 높았고, 10월이 가장 낮았다. 그리고 6, 7, 9월의 heat pulse 속도(速度)는 거의 비슷한 값을 나타냈다. 6. 줄기에 있어 방위별 heat pulse 속도(速度)는 동측이 가장 높았고, 서측과 북측은 비슷한 속도를 나타냈으며, 남측의 속도가 가장 낮았다. 7. 변재부(邊材部)에 있어서 깊이별 heat pulse 속도(速度)의 차이는 수피(樹皮)로부터 2cm 깊이에서 가장 높고, 다음이 1cm 깊이, 그리고 3cm 깊이에서 가장 낮았다. 8. 줄기에 있어 수분이동방향(水分移動方向)은 5본 모두 오른쪽 나선상승(螺旋上昇)(spiral ascent turning right)을 나타내고 있었다. 특히 지상의 3m까지는 매우 느린 회선(回旋)을 나타내다가 그 이상의 수고(樹高)에서는 매우 빠른 회선(回旋)으로 상승됨을 알 수 있었다. 9. 수액유량(樹液流量)(SF)은 SF=1.37AV 식(式)으로 나타났고, 이 식으로 구한 수액유량(樹液流量)은 우세목(優勢木)이 준우세목(準優勢木)과 열세목(劣勢木)보다 현저히 높았다. 10. 1ha의 임분(林分)에 대한 1일의 증산량(蒸散量)의 구성비율(構成比率)은 낮이 83%, 밤이 17%였고, 증산량(蒸散量)은 약 30.8ton/ha/day이었다. 11. 1ha의 월별(月別) 증산량(蒸散量)은 8월이 1,194ton/ha/month로 가장 많았고, 5월이 386ton/ha/month로 가장 낮았다. 또 1ha의 연간(年間) 증산량(蒸散量)은 3,983ton/ha/yr이었다. This is the basic study in order to know the amount of transpirational water loss in a Larix leptorepis stand by a heat pulse method. Especially this study has been measured and discussed the diurnal and seasonal trends of heat pulse velocity by changes of radiation, temperature and humidity, differences of heat pulse velocity by direction and depth in stem, differences of heat pulse velocity by dominant, codominant and suppressed trees, diurnal change of heat pulse velocity by change of leaf water potential, sap flow path way in sapwood by dye penetration and amount of daily and annual transpiration in a tree and stand. The results obtained as follows : 1. Relation between heat pulse velocity(V) and sap flow rate(SFR) was established as a equation of SFR=1.37V($r=0.96^{**}$). 2. The sap flow rate presented in the order of dominant, codominant and suppressed tree, respectively. The daily heat pulse velocity was changed by radiation, temperature and vapor pressure deficit. 3. The heat pulse velocity in individual trees did not differ in early morning and in late night, but had some differed from 12 to 16 hours when radiation was relatively high. 4. The heat pulse velocity and leaf water potential showed similar diurnal variation. 5. The seasonal variation of heat pulse velocity was highest in August, but lowest in October and similar value of heat pulse velocity in the other months. 6. The heat pulse velocity in stem by direction was highest in eastern, but lowest in southern and similar velocity in western and northern. 7. The difference of heat pulse velocity in according to depths was highest in 2.0cm depth, medium in 1.0cm depth, and lowest in 3.0cm depth from surface of stem. 8. The sap flow path way in stem showed spiral ascent turning right pattern in five sample trees, especially showed little spiral ascent turning right in lower part than 3m hight above ground, but very speedy in higher than 3m hight. 9. The amount of sap flow(SF) was presented as a equation of SF=1.37AV and especially SF in dominant tree was larger than in codominant or suppressed tree. 10. The amount of daily transpiration was 30.8ton/ha/day and its composition ratio was 83% at day and 17% at night. 11. The amount of stand transpiration per month was largest in August(1,194ton/ha/month), lowest in May (386ton/ha/month). The amount of stand transpiration per year was 3,983ton/ha/year.