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      • KCI등재

        Grazing Effects on Floristic Composition and Above Ground Plant Biomass of the Grasslands in the Northeastern Mongolian Steppes

        Hayashi, Ichiroku*,Kiyokazu Kawada,Mayu Kurosu,Amgaa Batjargal,Tsagaanbandi Tsundeekhuu,Toru Nakamura 한국생태학회 2008 Journal of Ecology and Environment Vol.31 No.2

        We describe plant biomass in the grasslands of the Mongolian steppe obtained using a quadrat sampling technique. Four sites were studied in the northeastern Mongolia located between 47°12'N and 47°40'N and 102°22'E and 112°24'E, which were typical grasslands of the steppe. Biomass, carbon and nitrogen content were determined for the plants collected from the grazed and ungarazed stands. With the measurements above, we expect to obtain information on grazing effects on the grasslands and carbon sequestration of the grassland from the air. In order to estimate the biomass without destroying the stands, we derived an equation to describe the relationship between plant biomass and v-value using plant height and species coverage within the stand. Estimated plant biomass in the ungrazed and grazed stands ranged between 108.0 g m-2 and 13.4 g m-2 and between 97.5 g m-2 and 14.1 g m-2 in late June 2005, respectively. Litter in the ungrazed and grazed stands ranged from 330.3 g m-2 to 78.4 g m-2 and from 188.0 g m-2 to 20.3 g m-2, similarly. Average carbon and nitrogen contents in plants and in litter were 43.0% and 1.9% and 33.7% and 1.4%, respectively. In study sites at Baganuur, the carbon and nitrogen content of plant materials (plant plus litter) was 118.4 g m-2 and 4.7 g m-2 on 30 June 2005. We describe plant biomass in the grasslands of the Mongolian steppe obtained using a quadrat sampling technique. Four sites were studied in the northeastern Mongolia located between 47°12'N and 47°40'N and 102°22'E and 112°24'E, which were typical grasslands of the steppe. Biomass, carbon and nitrogen content were determined for the plants collected from the grazed and ungarazed stands. With the measurements above, we expect to obtain information on grazing effects on the grasslands and carbon sequestration of the grassland from the air. In order to estimate the biomass without destroying the stands, we derived an equation to describe the relationship between plant biomass and v-value using plant height and species coverage within the stand. Estimated plant biomass in the ungrazed and grazed stands ranged between 108.0 g m-2 and 13.4 g m-2 and between 97.5 g m-2 and 14.1 g m-2 in late June 2005, respectively. Litter in the ungrazed and grazed stands ranged from 330.3 g m-2 to 78.4 g m-2 and from 188.0 g m-2 to 20.3 g m-2, similarly. Average carbon and nitrogen contents in plants and in litter were 43.0% and 1.9% and 33.7% and 1.4%, respectively. In study sites at Baganuur, the carbon and nitrogen content of plant materials (plant plus litter) was 118.4 g m-2 and 4.7 g m-2 on 30 June 2005.

      • KCI등재

        Phenological Studies of Deciduous Trees in the Cool Temperate Region of Japan

        Kato, Jun,Ichiroku Hayashi 한국생태학회 2008 Journal of Ecology and Environment Vol.31 No.3

        We obtained quantitative information on leaf unfolding and leaf shedding by observing 45 species of cool temperate deciduous trees in an arboretum over 5 growing seasons. These trees were in leaf (the foliage period) for 207 days on average after 1 April; 50% of leaves had been shed by 192 days after 1 April. Duration from the start of leaf unfolding to 50% leaf shedding was 157 days on average. Leaf unfolding began 35 days on average after 1 April. For leaf unfolding to begin, a 51℃․day of cumulated daily mean air temperature above 5℃ from 1 January (modified Kira's warmth index) was needed. Fifty-nine days elapsed between initiation and the final stage of leaf unfolding. The period of net photosynthetic assimilation was 157 days. The species with succeeding- type leaf unfolding associated with the anemochore seed type dominated the early stage of succession, while the species with flush-type leaf unfolding tended to dominate the late stage of succession. Few species were found in regions where late frosts occur after the day when the cumulative temperature for leaf unfolding is achieved. Biological characteristics include time of leaf unfolding, which affects the life history of each species, so that each species occupies its own niche in the stand. We conclude that that leaf phenology, such as timing of leaf unfolding and leaf shedding, is one of the components of each species' ecological characteristics. We obtained quantitative information on leaf unfolding and leaf shedding by observing 45 species of cool temperate deciduous trees in an arboretum over 5 growing seasons. These trees were in leaf (the foliage period) for 207 days on average after 1 April; 50% of leaves had been shed by 192 days after 1 April. Duration from the start of leaf unfolding to 50% leaf shedding was 157 days on average. Leaf unfolding began 35 days on average after 1 April. For leaf unfolding to begin, a 51℃․day of cumulated daily mean air temperature above 5℃ from 1 January (modified Kira's warmth index) was needed. Fifty-nine days elapsed between initiation and the final stage of leaf unfolding. The period of net photosynthetic assimilation was 157 days. The species with succeeding- type leaf unfolding associated with the anemochore seed type dominated the early stage of succession, while the species with flush-type leaf unfolding tended to dominate the late stage of succession. Few species were found in regions where late frosts occur after the day when the cumulative temperature for leaf unfolding is achieved. Biological characteristics include time of leaf unfolding, which affects the life history of each species, so that each species occupies its own niche in the stand. We conclude that that leaf phenology, such as timing of leaf unfolding and leaf shedding, is one of the components of each species' ecological characteristics.

      • SCOPUSKCI등재

        The Determination and Prediction of Pine to Oak Forest Succession in Sugadaira, Central Japan

        Jun, Kato,Hayashi, Ichiroku The Ecological Society of Korea 2003 Journal of Ecology and Environment Vol.26 No.4

        In order to analyze the succession process from a pine forest to an oak forest, the tree growth of Pinus densiflora and Quercus mongolica ssp. crispula was monitored in a permanent quadrat for 23 years. The measurements were carried out for the stem diameter (DBH) of Pinus densiflora between 1977 and 1999 and for the height of Quercus mongolica ssp. crispula saplings between 1998 and 2000. The floristic composition and the locations of the individual P. densiflora and Q. mongolica ssp. crispula trees and saplings in the quadrat were recorded. P densiflora and Q. mongolica ssp. crispula individuals were randomly distributed within the quadrat. The relative growth rates (RGR) of DBH in P. densiflora were 0.085 $yr^{-1}$ for large trees and 0.056 $yr^{-1}$ for small trees in 1977. The RGR of height for Q. mongolica ssp. crispula was 0.122 $yr^{-1}$. The growth curve for DBH of P. densiflora was approximated by the logistic equation: $$DBH(t) = 30 {[1+1.16exp(-0.13 t)]}^{-1}$$ where DBH (t) the DBH (cm) in year t and t is the number of years since 1977. The growth in height of P. densiflora and Q. mongolica ssp. crispula was described by following equations: $$H (t) = 20.2 {[1+0.407exp(-0.137 t)]}^{-1} (P. densiflora)$$ $$H (t) = 30 {[1+20.7exp(-0.122 t)}^{-1} (Q. mongolica ssp. crispula)$$ Where H (t) is the tree height (m) in year t and t is the number of years since 1977 in P. densiflora and 1998 in Q. mongolica ssp. crispula. With these equations we predicted that the height of Q. mongolica ssp. crispula increases from 2 m in 1999 to 20 m in 2029. Therefore, Q. mongolica ssp. crispula and P. densiflora will be approximately the same height in 2029. The years required for succession from a pine forest to an oak forest are expected 33 with the range between 23 and 44 years.

      • SCOPUSKCI등재

        Phenological Studies of Deciduous Trees in the Cool Temperate Region of Japan

        Jun, Kala,Hayashi, Ichiroku The Ecological Society of Korea 2008 Journal of Ecology and Environment Vol.31 No.3

        We obtained quantitative information on leaf unfolding and leaf shedding by observing 45 species of cool temperate deciduous trees in an arboretum over 5 growing seasons. These trees were in leaf (the foliage period) for 207 days on average after 1 April; 50% of leaves had been shed by 192 days after 1 April. Duration from the start of leaf unfolding to 50% leaf shedding was 157 days on average. Leaf unfolding began 35 days on average after 1 April. For leaf unfolding to begin, a$ 51^{\circ}C{\cdot}day$ of cumulated daily mean air temperature above $5^{\circ}C$ from 1 January (modified Kira's warmth index) was needed. Fifty-nine days elapsed between initiation and the final stage of leaf unfolding. The period of net photosynthetic assimilation was 157 days. The species with succeeding- type leaf unfolding associated with the anemochore seed type dominated the early stage of succession, while the species with flush-type leaf unfolding tended to dominate the late stage of succession. Few species were found in regions where late frosts occur after the day when the cumulative temperature for leaf unfolding is achieved. Biological characteristics include time of leaf unfolding, which affects the life history of each species, so that each species occupies its own niche in the stand. We conclude that that leaf phenology, such as timing of leaf unfolding and leaf shedding, is one of the components of each species' ecological characteristics.

      • KCI등재

        Factors affecting the vertical distribution of Betula platyphylla var. japonica and Betula ermanii on Mt. Neko in Nagano Prefecture, Japan

        Otsubo Jiro,Mariko Shigeru,Hayashi Ichiroku 한국생태학회 2010 Journal of Ecology and Environment Vol.33 No.2

        Betula platyphylla var. japonica and Betula ermanii segregate vertically at an elevation of approximately 1,850 m on Mt. Neko in Nagano Prefecture, Japan. B. platyphylla var. japonica and B. ermanii were the dominant species below and above this altitude, at which the mean-annual and growing-season air temperatures were 4°C and 14.1°C, respectively. Based on a modification of Kira's warmth index which employs cumulative temperature represented as °C day, leaf unfolding in both species was observed to be initiated at 58°C day and 169°C day, respectively. In 1996, leaf unfolding was initiated on 18 May in B. platyphylla var. japonica (+/-6 days) and on 5 June in B. ermanii (+/-8 days), shortly after the last frost which occurred on 5 May 1995 above 1,850 m; below this elevation there was no risk of frost at the time. At elevations above 1,850 m, the unfolded leaves of B. platyphylla were damaged by late frost, while B. ermanii escaped injury because the leaves were still protected by winter buds. The optimum temperature for seed germination in both B. platyphylla and B. ermanii was 30°C. Temperature alternation from 10 to 30°C and moist storage of seeds at 4°C (stratification) prior to incubation increased germination rates in both species. The seedlings of B. ermanii had a greater survival rates than those of B. platyphylla var. japonica when planted above 1,850 m. Comparisons of the timing of leaf unfolding and the latest frost at a site appeared to be the main factors affecting the vertical distribution of these species.

      • SCOPUSKCI등재

        Factors affecting the vertical distribution of Betula platyphylla var. japonica and Betula ermanii on Mt. Neko in Nagano Prefecture, Japan

        Jiro, Otsubo,Shigeru, Mariko,Ichiroku, Hayashi The Ecological Society of Korea 2010 Journal of Ecology and Environment Vol.33 No.2

        Betula platyphylla var. japonica and Betula ermanii segregate vertically at an elevation of approximately 1,850 m on Mt. Neko in Nagano Prefecture, Japan. B. platyphylla var. japonica and B. ermanii were the dominant species below and above this altitude, at which the mean-annual and growing-season air temperatures were $4^{\circ}C$ and $14.1^{\circ}C$, respectively. Based on a modification of Kira's warmth index which employs cumulative temperature represented as $^{\circ}C$ day, leaf unfolding in both species was observed to be initiated at $58^{\circ}C$ day and $169^{\circ}C$ day, respectively. In 1996, leaf unfolding was initiated on 18 May in B. platyphylla var. japonica (+/-6 days) and on 5 June in B. ermanii (+/-8 days), shortly after the last frost which occurred on 5 May 1995 above 1,850 m; below this elevation there was no risk of frost at the time. At elevations above 1,850 m, the unfolded leaves of B. platyphylla were damaged by late frost, while B. ermanii escaped injury because the leaves were still protected by winter buds. The optimum temperature for seed germination in both B. platyphylla and B. ermanii was $30^{\circ}C$. Temperature alternation from 10 to $30^{\circ}C$ and moist storage of seeds at $4^{\circ}C$ (stratification) prior to incubation increased germination rates in both species. The seedlings of B. ermanii had a greater survival rates than those of B. platyphylla var. japonica when planted above 1,850 m. Comparisons of the timing of leaf unfolding and the latest frost at a site appeared to be the main factors affecting the vertical distribution of these species.

      • KCI등재

        Floristic Composition, Grazing Effects and Above-ground Plant Biomass in the Hulunbeier Grasslands of Inner Mongolia, China

        Kawada, Kiyokazu,Mayu Kurosu,Yunxiang Cheng,Tsagaanbandi Tsendeekhuu,Wuyunna,Toru Nakamura,Ichiroku Hayashi 한국생태학회 2008 Journal of Ecology and Environment Vol.31 No.4

        In order to assess the effect of grazing on grasslands using floristic composition, we studied the floristic composition and plant biomass of steppe vegetation in Hulunbeier, Inner Mongolia. The dominant species of the grasslands were Stipa krylovii, Stipa grandis and Leymus chinensis. Floristic composition changed according to grazing pressure, being light, heavy and moderate. Under heavy grazing conditions, the importance value of P. acaulis, C. korshinskyi and Cleistogenes squarrosa increased in the plant community. These species were indicators of heavily grazed stand. Plant biomass of the stands ranged within 11.6 g m-2 and 63.5 g m-² and 69.5 g m-² and 166.2 g m-² to the west and east of Lake Hulun, respectively. These values are equivalent to ca. 450~1,000 kg of atmospheric carbon per hector, which is retained within the stand during the summer season. In order to assess the effect of grazing on grasslands using floristic composition, we studied the floristic composition and plant biomass of steppe vegetation in Hulunbeier, Inner Mongolia. The dominant species of the grasslands were Stipa krylovii, Stipa grandis and Leymus chinensis. Floristic composition changed according to grazing pressure, being light, heavy and moderate. Under heavy grazing conditions, the importance value of P. acaulis, C. korshinskyi and Cleistogenes squarrosa increased in the plant community. These species were indicators of heavily grazed stand. Plant biomass of the stands ranged within 11.6 g m-2 and 63.5 g m-² and 69.5 g m-² and 166.2 g m-² to the west and east of Lake Hulun, respectively. These values are equivalent to ca. 450~1,000 kg of atmospheric carbon per hector, which is retained within the stand during the summer season.

      • KCI등재

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