1 이일로, "미토콘드리아 COI와 핵 RAG1 유전자 분석에 의한 줄종개(Cobitis tetralineata)와 왕종개(Iksookimia longicorpa) 간 자연잡종 동정" 한국어류학회 21 (21): 287-290, 2010
2 김치홍, "각시붕어 Rhodeus uyekii와 묵납자루 Acheilognathus signifer (Pisces; Cyprinidae)의 속간 자연 잡종의 출현" 한국어류학회 22 (22): 225-229, 2010
3 Gerber, A. S., "The role of introgressive hybridization in the evolution of the Gila robusta complex(Teleostei : Cyprinidae)" 55 : 2028-2039, 2001
4 Allendorf, F. W., "The problems with hybrids : setting conservation guidelines" 16 : 613-622, 2001
5 Damme, V. D., "The introduction of the European bitterling(Rhodeus amarus)to west and central Europe" 8 : 79-106, 2007
6 Kim, D. M., "Tanakia latimarginata, a new species of bitterling from the Nakdong River, South Korea(Teleostei : Cyprinidae)" 25 : 59-68, 2014
7 윤영은, "RAG-1 유전자의 염기서열 분석에 의한 각시붕어 Rhodeus uyekii와 떡납줄갱이 R. notatus 잡종의 동정" 한국양식학회 22 (22): 79-82, 2009
8 Campton, D. E, "Population Genetics and Fishery Management" University of Washington Press 161-192, 1987
9 Okazaki, M., "Phylogenetic relationships of bitterlings based on mitochondrial 12S ribosomal DNA sequences" 58 : 89-106, 2001
10 김형수, "Occurrence of a Natural Intergeneric Hybrid between Rhodeus pseudosericeus and Acheilognathus signifer (Pisces: Cyprinidae) from the Namhangang (river), Korea" 한국어류학회 26 (26): 153-158, 2014
1 이일로, "미토콘드리아 COI와 핵 RAG1 유전자 분석에 의한 줄종개(Cobitis tetralineata)와 왕종개(Iksookimia longicorpa) 간 자연잡종 동정" 한국어류학회 21 (21): 287-290, 2010
2 김치홍, "각시붕어 Rhodeus uyekii와 묵납자루 Acheilognathus signifer (Pisces; Cyprinidae)의 속간 자연 잡종의 출현" 한국어류학회 22 (22): 225-229, 2010
3 Gerber, A. S., "The role of introgressive hybridization in the evolution of the Gila robusta complex(Teleostei : Cyprinidae)" 55 : 2028-2039, 2001
4 Allendorf, F. W., "The problems with hybrids : setting conservation guidelines" 16 : 613-622, 2001
5 Damme, V. D., "The introduction of the European bitterling(Rhodeus amarus)to west and central Europe" 8 : 79-106, 2007
6 Kim, D. M., "Tanakia latimarginata, a new species of bitterling from the Nakdong River, South Korea(Teleostei : Cyprinidae)" 25 : 59-68, 2014
7 윤영은, "RAG-1 유전자의 염기서열 분석에 의한 각시붕어 Rhodeus uyekii와 떡납줄갱이 R. notatus 잡종의 동정" 한국양식학회 22 (22): 79-82, 2009
8 Campton, D. E, "Population Genetics and Fishery Management" University of Washington Press 161-192, 1987
9 Okazaki, M., "Phylogenetic relationships of bitterlings based on mitochondrial 12S ribosomal DNA sequences" 58 : 89-106, 2001
10 김형수, "Occurrence of a Natural Intergeneric Hybrid between Rhodeus pseudosericeus and Acheilognathus signifer (Pisces: Cyprinidae) from the Namhangang (river), Korea" 한국어류학회 26 (26): 153-158, 2014
11 Kim, I. S, "Illustrated encyclopedia of fauna and flora of Korea. VoL. 37. Freshwater fishes"
12 Grant, P. R., "Hybridization of bird species" 256 : 193-197, 1992
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14 Avise, J. C., "Hybridization and introgression among species of sunfish(Lepomis) : analysis by mitochondrial DNA and allozyme markers" 108 : 237-255, 1984
15 Jansson, H., "High frequency of natural hybrids between Atlantic salmon, Salmo salar L., and brown trout, S. trutta L., in a Swedish river" 39 : 343-348, 1991
16 Kim, I. S., "Freshwater fishes of Korea" Kyohak publishing 466-, 2002
17 Nelson, J. S, "Fishes of the world (4th ed)" John Wiley and Sons, Inc 601-, 2006
18 Hubbs, C. L., "Fishes of the great lakes region" The Univ. Michigan Press 250-, 2004
19 Nikoljukin, M. J, "Distant hybridization in acipenseridae and teleostei" Moskava 335-, 1972
20 Kim, I. S., "Diploid-triploid hybrid complex of the spined loach Cobitis sinensis and C. longicorpus(Pisces, Cobitidae)" 2 : 203-210, 1990
21 Kim, I. S., "An occurrence of intergeneric hybrid cross, Pungtungia herzi×Pseudopungtungia nigra from the Ungcheon River, Korea" 3 : 42-47, 1991
22 Sonnenberg, R., "An evaluation of LSU rDNA D1-D2 sequences for their use in species identification" 4 : 6-, 2007
23 Šorić V. M, "A natural hybrid of Leuciscus cephalus and Alburnus alburnus(Pisces, Cyprinidae)from the Ibar River(Western Serbia)" 56 : 23-32, 2004
24 Hwang, Y. J., "A natural hybrid between spinous loach, Cobitis longicorpus and cyprinid loach Misgurunus anguilicaudatus(Pisces, Cobitidae)" 7 : 203-207, 1995
25 김근용, "A Natural Hybrid of Intergeneric Mating between a Female Pungtungia herzi and a Male Pseudorasbora parva (Cypriniformes: Cyprinidae)" 한국수산과학회 18 (18): 99-107, 2015