1 Cambefort Y., "Étude écologique des Coléoptères Scarabaeidae de Côte d’Ivoire" 3 (3): 1-294, 1984
2 Janssens A, "qContributions à l’étude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). TroisièmePartie. XC. Coleoptera Scarabaeidae. Scarabaeinae" 51 : 343-356, 1956
3 Kraatz G., "Zwei neue westafrikanische Oniticellus-Arten" 34 (34): 142-144, 1895
4 Kolbe HJ, "Wissenschaftliche Ergebnisse der deutschen Zentral Afrika Expedition 1907-1908, Band V, Lieferung 3. Klinkhardt and Biermann" 261-416, 1914
5 d’Orbigny H, "Voyage dans l’Afrique orientale anglaise 1912-1913" Résultats scientifiques 1-31, 1916
6 Harold Ev., "Verzeichniss der von Dr. Beccari in Bogos gesammelten coprophagen Lamellicornien" 8 : 1-28, 1871
7 Cambefort Y, "The nesting behavior of dung beetles (Scarabaeinae), an ecological and evolutive approach" Publicaciones del Instituto de Ecologia 1-176, 1982
8 Arrow GJ, "The fauna of British India including Ceylon & Burma, Part 3, Coleoptera:Lamellicornia (Coprinae)" Taylor and Francis 1-353, 1931
9 Balthasar V, "The Scientific Results of the Hungarian Soil Zoological Expedition to the Brazzaville-Congo. 22. Scarabaeinae und Coprinae(Coleoptera). (131. Beitrag zur Kenntnis der Scarabaeoidea)" 7 : 47-73, 1967
10 Davis ALV, "The African Dung Beetle Genera" Protea Book House 1-272, 2008
11 Chandra K, "Taxonomic studies on dung beetles(Coleoptera : Scarabaeidae, Geotrupidae, Hybosoridae)of Chhattisgarh, India" 8 (8): 331-360, 2013
12 Boucomont A., "Synopsis des"Oniticellini"d’Afrique" 9 : 197-234, 1921
13 Sabu TK, "Succession of dung beetles (Scarabaeinae:Coleoptera) in the dung pats of gaur, Bos gaurus H. Smith (Artiodactyla: Bovidae), from the moist deciduous forests of southern Western Ghats" 1 (1): 59-69, 2007
14 Ochi T, "Studies on the Coprophagous Scarab Beetles from East Asia. V (Coleoptera, Scarabaeidae)" 8 : 29-35, 1996
15 Vinod KV, "Species composition and community structure of dung beetles attracted to dung of gaur and elephant in the moist forests of South Western Ghats" 7 (7): 1-14, 2007
16 Ferreira MC, "South African Animal Life. Results of the Lund University Expedition in 1950-1951" Almqvist V, Wiksells Boktryckeri AB 479-516, 1958
17 Cambefort Y., "Scarabaeinae récoltés dans le Fouta Djalon(Guinée)" 33 (33): 420-424, 1971
18 Boucomont A, "Scarabaeidae: Coprinae, Termitotroginae. Partes 38 et 90" W. Junk 103-263, 1927
19 Balthasar V, "Scarabaeidae des paläarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Bestimmungs-Tabellen der europäischen Coleopteren. Troppau:115. Heft" Verlag E. Reitter 1-112, 1935
20 Chandra K, "Scarab beetles(Coleoptera : Scarabaeoidea)of Barnawapara Wildlife Sanctuary, Chhattisgarh, India" 5 (5): 4660-4671, 2013
21 Branco T., "Revision of the genus Tiniocellus Péringuey, 1901 and Nitiocellus gen. n. (Coleoptera, Scarabaeidae, Oniticellini)" 71-126, 1901
22 Emlen DJ, "Phylogenetic evidence for an association between tunnelling behavior and the evolution of horns in dung beetles(Coleoptera : Scarabaeidae : Scarabaeinae)" 5 : 47-56, 2006
23 Ferreira MC., "Os Escarabídeos de África (sul do Sáara). I" 11 : 1-1088, 1972
24 Ferreira MC., "Os Escarabídeos de Moçambique. I (Subfamílias Scarabaeinae Coprinae)" 10 : 5-778, 1968
25 Ferreira MC, "Os Escarabídeos da sub-região natural da África do Sul. 9. Chaves para determinação das espécies dos generous Drepanocerus Kirby, Euoniticellus Janssens, Tiniocellus Péringuey, Liatongus Reittere, Oniticellus Serville (Tribo Oniticellini)" 9 (9): 39-57, 1967
26 Balthasar V, "Oniticellini, eineneue Art von Drepanocerus. 113. Beitragzur Kenntnis der Scarabaeoidea, Col. Parc National de la Garamba - Mission H. de Saeger en collaborationavec P. Baert, G. Demoulin, I. Denisoff, J. Martin, M. Micha, A. Noirfalise, P. Schoemaker, G. Troupin et J. Verschuren (1941-1952)" 40 (40): 131-135, 1963
27 Sarkar SK, "On the taxonomy of Scarabaeine fauna(Coleoptra : Scarabaeidae)of Buxa Tiger Reserve(BTR), West Bengal, India" 10 (10): 18-48, 2015
28 Arrow GJ., "Notes on the Coleopterous Genus Oniticellus and descriptions of some new species from India" 1 (1): 178-183, 1908
29 Cambefort Y, "Nidification et larves des Oniticellini afro-tropicaux (Col. Scarabaeidae). (Comptes Rendues du 1er Congres International des Entomologistes d’expression française, Paris, 6-9 juill. 1982)" 88 : 542-569, 1983
30 Balthasar V, "Neue Scarabaeiden-Arten. 129. Beitrag zur Kenntnis der Scarabaeoidea(Coleoptera)" 18 : 953-958, 1968
31 Balthasar V, "Monographie der Scarabeidae und Aphodiidae der palaearktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2. Coprinae (Onitini, Oniticellini, Onthophagini)" Verlag der Tschechoslowakischen Akademieder Wissenschaften 1-627, 1963
32 Ferreira MC, "Monografia dos Escarabídeos da África do Sul. 3 Parte. Tribo Oniticellini. No 2 - Géneros Oniticellus Serv., Euoniticellus A. Janssens, Liatongus Reitter, Tiniocellus Péringuey - Tragiscus Klug" 91 : 81-123, 1955
33 Müller G, "Missione biologica nel paese dei Borana. Raccolte zoologiche. Vol. II, Pt. I" Reale Accademia d’Italia 1-150, 1940
34 Balthasar V, "Mission Zoologique de l’I.R.S.A.C. en Afrique Orientale (P. Basilewsky et N. Leleup, 1957). Résultats Scientifiques. Deuxième Partie. 41. Coleoptera Scarabaeidae: Scarabaeini, Coprini, Onitini et Oniticellini. (101. Beitrag zur Kenntnis der Scarabaeiden)" 88 : 55-66, 1960
35 Davis ALV, "Methods for the inventory and ecological monitoring of dung beetles, butterflies and termites in the East Usambaras" 35-46, 1996
36 Cambefort Y., "Les Scarabaeidae s. str. de Lamto (Côte d’Ivoire): structure des peuplements et rôle dans l’écosystème" 18 : 433-459, 1982
37 Cambefort Y., "Les Coléoptères Scarabaeidae du Parc National de Taï(Côte d’Ivoire)" 7 : 337-342, 1986
38 Endrödi S., "Lamellicornia(Coleoptera)aus Ghana, II" 68 : 155-164, 1976
39 Fåhraeus OI, "Insecta Caffrariae annis 1838-1845 a J. A. Wahlberg collecta. Pars 2, Coleoptera (Scarabaeides)" Officina Norstedtiana 1-396, 1857
40 Davis ALV., "Habitat associations in a South African, summer rainfall, dung beetle community(Coleoptera : Scarabaeidae, Aphodiidae, Staphylinidae, Histeridae, Hydrophilidae)" 40 : 260-280, 1996
41 Sabu TK, "Guild structure, diversity and succession of dung beetles associated with Indian Elephant dung in South Western Ghats forests" 6 (6): 1-12, 2006
42 Singh AP, "First report on Scarabaeid(Coleoptera)fauna of Sahaspur, Uttarakhand" 5 (5): 17-22, 2017
43 Sabu TK, "Faunal survey, endemism and possible species loss of Scarabaeinae(Coleoptera : Scarabaeidae)in the western slopes of the moist South Western Ghats, South India" 2830 : 29-38, 2011
44 Ferreira MC, "Exploration du Parc national de la Garamba, Mission H. de Saeger. Faciscule 30: Coprinae (Coleoptera Lamellicornia). Fam. Scarabaeidae" Institut des Parcs Nationauxdu Congo et du Rwanda 1-123, 1962
45 Janssens A., "Exploration du Parc National de l’Upemba" Institut des Parcs Nationaux du Congo Belge 1-118, 1953
46 Gestro R, "Esplorazione del Giuba e dei suoi affluent compiuta dal Cap. V. Bottego durante gli anni 1892-93 sotto gli auspicii della Società Geografica italiana. Risultati zoologici. XVI. Coleotteri" 15 : 247-494, 1895
47 Endrödi S, "Entomological Explorations in Ghana by Dr. S. Endrödy-Younga. 14. Lamellicornia (Coloeoptera), I" 65 : 195-229, 1973
48 Botes A, "Elephant-and human-induced changes to dung beetles(Coleopetra : Scarabaeidae)assemblages in the Maputa land centre of endemism" 130 : 573-583, 2006
49 Davis ALV, "Effect of deforestation on a southwest Ghana dung beetle assemblage(Coleoptera : Scarabaeidae)at the periphery of Ankasa conservation area" 34 : 1081-1088, 2005
50 Cambefort Y, "Dung Beetle Ecology" Princeton University Press 1-481, 1991
51 Hanski I, "Dung Beetle Ecology" Princeton University Press 1-481, 1991
52 Cambefort Y, "Dung Beetle Ecology" Princeton University Press 156-178, 1991
53 Cambefort Y., "Données préliminaires sur l’écologie des Scarabaeinae coprophages de Lamto(Insecta, Coleoptera, Scarabaeoidea)" 13 : 61-79, 1980
54 Davis ALV., "Diel and seasonal community dynamics in an assemblage of coprophagous, Afrotropical, dung beetles (Coleoptera: Scarabaeidae s. str., Aphodiidae, and Staphylinidae: Oxytelinae)" 110 : 291-308, 1996
55 Gerstaecker A, "Die Gliederthier-Fauna des Sansibar- Gebietes. Nach dem von Dr. O. Kersten während der v. d. Decken’schen Ost-Afrikanischen Expedition im Jahre 1862 und von C. Cooke auf der Insel Sansibarim Jahre 1864 gesammelten Material" C.F. Winter 1-544, 1873
56 Roth JR., "Diagnosen neuer Coleoptera aus Abyssinien" 17 (17): 115-133, 1851
57 Péringuey L., "Descriptive catalogue of the Coleoptera of South Africa(Lucanidae and Scarabaeidae)" 12 (12): 1-563, 1901
58 Gillet JJE, "Coprophages et Dynastides provenant du voyage de M. F. Colmant dans les régions de l’Uellé, du Bomu et du Bahr el Ghazal" 52 : 54-67, 1908
59 Boucomont A., "Coprophaga africana, 2e note" 11 : 40-58, 1923
60 Janssens A, "Coprini. Coleoptera Lamellicornia, Fam. Scarabaeidae" Exploration du Parc national Albert, Mission G. F. de Witte 1-103, 1939
61 Balthasar V, "Contributions à la connaissance de la Faune Entomologique de la Côte-d’Ivoire. Deuxième Partie: Annales du Musée Royal de l’Afrique Centrale, Série in-8o - Sciences Zoologiques" 1-100, 1969
62 Durand H., "Contribution à l’étudebiologique du Sénégal septentrional. XVI. Coléoptères Scarabaeidae Scarabaeinae et Chironidae" 34 : 78-90, 1972
63 Paulian R, "Contribution à l’étude de la faune entomologique de l’Angola. Mission scientifique Suisse en Angola. Coléoptères Lamellicornes Laparosticti" 105 : 1-41, 1937
64 Kabakov ON, "Contribution to the knowledge of the subfamily Scarabaeinae from China with nomenclatural notes" 10 (10): 47-59, 2014
65 Barbero EL, "Contribution to the knowledge of Scarabaeidae and Aphodiidae from southern Ethiopia(Coleoptera : Scarabaeoidea)" 130 : 233-254, 1998
66 Ferreira MC., "Contribuição para o conhecimento dos Escarabídeos de Angola" 54 : 121-190, 1962
67 Nel A, "Comparative morphology of the mouthparts of adult Scarabaeoidea, vol. 80. Entomology Memoir" Department of Agricultural Development Republic of South Africa 1-81, 1990
68 Davis ALV., "Community organization of dung beetles(Coleoptera : Scarabaeidae) : differences in body size and functional group structure between habitats" 34 : 258-275, 1996
69 Bates HW., "Coleoptera from Kulu in NW India" 24 : 7-23, 1891
70 Davis ALV., "Climatic and biogeographical associations of southern African dung beetles (Coleoptera: Scarabaeidae s. str.)" 35 : 10-38, 1997
71 Davis ALV, "Climatic and biogeographical associations of Kenyan and northern Tanzanian dung beetle(Coleoptera : Scarabaeidae)" 31 : 290-305, 1993
72 Krajcik M., "Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera:Scarabaeidae: Scarabaeinae). Anima. X, Supplement 3, Plzen" 1-189, 2006
73 Ferreira MC., "Catálogo dos Coleópteros de Angola" 8 : 417-1317, 1967
74 Harold Ev, "Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus, vol. 4" Gummi E.H. 979-1346, 1869
75 Bezdek A, "Catalogue of Palaearctic Coleoptera. Volume 3:ScarabaeoideaeScirtoidea Dascilloidea - Buprestoidea - Byrrhoidea" Apollo Books 1-690, 2006
76 Schoolmeesters P., "Catalogue of Life: Annual Checklist"
77 Gerstaecker A, "Beitrag zur Insekten-fauna von Zanzibar. III. Coleoptera(Fortsetzung von Jahrrg. XXXIII. p. 49)" 37 : 42-86, 1871
78 Davis ALV., "Associations of Afrotropical Coleoptera(Scarabaeidae : Aphodiidae : Staphylinidae : Hydrophilidae : Histeridae)with dung and decaying matter : implications for selection of fly-control agents for Australia" 28 : 383-399, 1994
79 Philips TK, "A phylogenetic analysis of dung beetles(Scarabaeinae : Scarabaeidae) : unrolling an evolutionary history" 18 : 53-88, 2004
80 Sarkar SK, "A first faunistic study on the tribe Oniticellini Kolbe, 1905(Coleoptera : Scarabaeidae)of Baikunthapur Tropical Forest of the Himalayan foothills, West Bengal, India" 8 : 1-19, 2020
81 Davis ALV, "A dung beetle survey of selected Gauteng nature reserves : implications for conservation of the provincial scarabaeini fauna" 13 : 1-16, 2005
82 Bacchus ME., "A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae)and the smaller Lamellicorn families (Coleoptera) described by G. J. Arrow" 37 (37): 97-115, 1978
83 Gajendra N, "A Reviewof Coleoptera diversity of Chhattisgarh : Updated checklist 2015" 5 (5): 710-714, 2016