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김계환(Kae - Hwan Kim),F . A . Aravanopoulos(F . A . Aravanopoulos),L . Zsuffa(L . Zsuffa) 한국산림과학회 1997 한국산림과학회지 Vol.86 No.2
The pollen morphology of 20 species and one variety of the two genera of Hippocastanaceae was investigated by LM, SEM and TEM. The pollen grains of this family are monads, prolate to subprolate, isopolar, radially symmetrical, 3-colporate or 3-colpate and ora is circular or lolongate. Sculpture pattern is distinctly striate or spinulose, Brae in various directions. Based on the aperture types and sculpture patterns three major pollen types of this family are discernible : hippocastanum, indica, and punduana.